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收稿日期:2019-07-25 修回日期:2019-08-22资助项目:国家自然科学基金(41771322); 国家重点研发计划(2017YFC0504504)第一作者:潘天辉(1993—),男,内蒙古阿拉善盟人,硕士研究生,研究方向:植物生理生态。E-mail:1406234356@qq.com通信作者:杜峰(1971—),男,内蒙古乌兰察布人,博士,副研究员,主要从事植物种群、群落生态及生理生态的研究。E-mail:dufeng@ms.iswc.ac.cn