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收稿日期:2017-07-06;改回日期:2017-07-25。
基金项目:国家重点研发计划(2016YFC0501802);青海省国际合作项目(2015-HZ-801)
作者简介:贺慧丹(1991-),女,山东菏泽人,博士研究生,主要从事全球变化生态学研究。E-mail:hhdan712@foxmail.com
通讯作者:李英年(1962-),男,青海乐都人,研究员,博士生导师,主要从事全球变化生态学研究。E-mail:ynli@nwipb.cas.cn