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收稿日期:2016-06-14;改回日期:2016-06-27。
基金项目:北京市农林科学院科技创新专项(KJCX20140419);北京市自然科学基金(6122010)
通讯作者:袁小环(1975-),女,安徽砀山人,博士,副研究员,主要从事园林植物生理生态研究。E-mail:sunringner@163.com