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收稿日期:2015-03-19;改回日期:2015-04-10。
基金项目:国家自然科学基金(31170575,41373075)
作者简介:裴忠雪(1990-),女(满),黑龙江省克山县人,硕士研究生,研究方向:植物生态学。E-mail:1009764815@qq.com
通讯作者:王文杰(1974-),男,河北省易县人,教授,研究方向:植物生态学。E-mail:wjwang225@hotmail.com