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收稿日期:2019-04-09 修回日期:2019-06-14 资助项目:国家自然科学基金(41563008); 云南省高原湿地科学创新团队(2012HC007) 第一作者:方昕(1994—),男,四川西昌人,硕士研究生,研究方向为湿地土壤微生物。E-mail:xcfangxin@163.com 通信作者:郭雪莲(1979—),女,吉林通榆人,副教授,硕士生导师,主要从事湿地生态学和生物地球化学研究。E-mail:guoxuelian2009@hotmail.com