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收稿日期:2018-12-18;改回日期:2018-12-29。
基金项目:国家自然科学基金重点项目(41730752);国家自然科学基金青年项目(31700395);青海省自然科学基金青年项目(2016-ZJ-918Q)
作者简介:郭小伟(1985-),男,山东潍坊人,博士,高级工程师,主要从事植物生态学研究。E-mail:xwguo1206@163.com
通讯作者:杜岩功(1981-)男,山东威海人,博士,副研究员,主要从事草地生态学研究。E-mail:ygdu@nwipb.cas.cn