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收稿日期:2016-07-30;改回日期:2016-08-12。
基金项目:国家自然科学基金项目(41301197);中国博士后科学基金资助项目(2014M550382,2015T80766);河南省高校科技创新团队支持计划(16IRTSTHN012)
作者简介:赵清贺(1982-),男,河南开封人,博士,讲师,主要从事流域景观格局与生态过程研究。E-mail:zqh410224@126.com